Operator analysis
Operator                                          Tuning   Count      Time     Time/Op  Pr(accept)  Performance suggestion
scale(ac)                                         0.715   139        746      5.37     0.2374      good	
scale(ag)                                         0.687   143        762      5.33     0.1469      good	
scale(at)                                         0.668   139        756      5.44     0.1727      good	
scale(cg)                                         0.622   117        641      5.48     0.265       good	
scale(gt)                                         0.611   116        618      5.33     0.3793      good	
frequencies                                       0.037   129        696      5.4      0.2403      good	
scale(alpha)                                      0.738   133        705      5.3      0.0827      slightly low	Try setting scaleFactor to about 0.8593
subtreeSlide(treeModel)                           0.029   19618      14961    0.76     0.0904      slightly low	Try decreasing size to about 0.014621959440898797
Narrow Exchange(treeModel)                                19592      13254    0.68     0.0191      low	
Wide Exchange(treeModel)                                  3824       2495     0.65     0.0016      low	
wilsonBalding(treeModel)                                  3943       4500     1.14     0.0041      low	
scale(treeModel.rootHeight)                       0.881   3991       1014     0.25     0.1185      good	
uniform(nodeHeights(treeModel))                           39189      35759    0.91     0.1298      good	
scale(yule.birthRate)                             0.512   4008       398      0.1      0.2872      good	
up:down:nodeHeights(treeModel)                    0.924   3919       21251    5.42     0.1638      good