('Ixiolaena',((Psychrophyton,(Pterygopappus,(Loricaria,(Sinoleontopodium,(Raouliopsis,Mniodes))))),(Antithrixia,(Oreoleysera,(Leysera,Rosenia,(Relhania,Oedera))),(Bryomorphe,(((Phaenocoma,Dolichothrix),Planea,Metalasia,Hydroidea,Atrichantha,Calotesta),(Lachnospermum,(Elytropappus,Stoebe,(Disparago,Amphiglossa))))))))