('Arctotidinae',(Cylindrocline,((Thespidium,(Coleocoma,Delamerea)),(((Stenachaenium,Stenachaenium_campestre),(Neojeffreya,(Pterocaulon,Monenteles))),((Pseudoconyza,(Blumeopsis,(Laggera,Nicolasia)),(Doellia,Porphyrostemma)),((Streptoglossa,Karelinia,((Monarrhenus,Berthelotia_lanceolata),Tecmarsis_bojeri,(Pluchea_odorata,Tessaria))),((Merrittia,(Eyrea_dentex,Adelostigma)),(Epaltes,(Litogyne,(Triplocephalum,(Sphaeranthus,Pseudoblepharispermum)))))))))))