(Ludwigia_peploides,((Oenothera_biennis,((Fuschia_x_triphylla,Hauya_heydeana),((Epilobium_ciliatum,Epilobium_hirsutum),((Epilobium_pallidiflorum,Epilobium_billardierianum,Epilobium_royleanum),Epilobium_melanocaulon)))),Circaea_cordata))