(Ludwigia_peploides,(((Fuschia_x_triphylla,Hauya_heydeana),(Oenothera_biennis,(((Epilobium_ciliatum,Epilobium_royleanum),Epilobium_hirsutum),((Epilobium_pallidiflorum,Epilobium_billardierianum),Epilobium_melanocaulon)))),Circaea_cordata))